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The article “a", in “but a gene could" (line 2), refers to a specific gene.
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Nowadays there are far more centenarians than expected.
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“Besides", in “Besides being at least 80" (line 7), and in addition to are synonyms.
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A synonymous word for “thus", in “It is thus widely believed" (lines 16 and 17), is therefore.
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According to the last paragraph, the discovery of the genes referred to by Eric Topol is only a question of time.